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Cardio
Conditioning - Applying the Science
THE
HEART AND THE CARDIOVASCULAR SYSTEM
Lesley Young, Centre
for Equine Studies, Animal Health Trust, Newmarket, UK.
Your horse's heart is
a pump that supplies all his muscles and organs with oxygen and
nutrients and carries away the waste products. It does this under all
conditions: when he is galloping after hounds, or when he's asleep.
To do this
efficiently, the heart is divided into two sides: the left side takes
oxygenated blood from the lungs and supplies muscles and vital organs,
like the brain and kidney. The de-oxygenated blood then returns to the
right heart to be pumped to the lungs, where it is replenished with
oxygen. Each side has a muscular pumping chamber, the ventricle, and a
collecting chamber, the atrium. With each beat, a typical adult horse
pumps approximately one litre of blood out of each side of his heart,
this is the stroke volume. With a resting heart rate of 30 - 40 beats
per minute, this means the total blood pumped in one minute, the horse's
cardiac output, is 30 - 40 litres per minute. Contrast this to man with
an average output of 5 litres per minute and we can begin to see why the
horse is a much better athlete.
How does the heart
supply the body under the stresses of exercise
The heart is an "intelligent" pump, but has a fairly limited
range of options to increase its output to meet the increasing demands
of exercise.
1: Increase the
amount of blood pumped with each beat; increase stroke volume
When the heart beats, it
does not empty completely, instead a proportion of blood remains in the
left heart chamber. When exercise begins, circulating adrenaline makes
the contraction of the heart muscle more forceful, so more blood is
ejected from the ventricle with each beat. Additionally the huge muscles
of locomotion contract and squeeze more blood back to the heart
increasing the amount of blood available in the circulation and further
increasing stroke volume.
2: Increase
heart rate:
Resting heart rate is
between 28 and 36 beats per minute in Thoroughbred horses. Maximum heart
rate during peak exercise in the Thoroughbred is between 225 and 240 per
minute. The huge range of heart rate is the most important mechanism for
increasing cardiac output during exercise, especially as work intensity
increases. Heart rates above 200 beats per minute ensure that cardiac
output increases from 35 litres per minute at rest to above 200 litres
per minute during maximal exercise.
What happens when
my horse gets fit?
Just as the muscles of locomotion become stronger and better able to do
their job efficiently, so too does the heart muscle. Using ultrasound,
(a method to visualise the horse's heart), we are now able to measure
cardiac dimensions very accurately and assess how efficiently it works.
We can also monitor what happens as the horse goes through its training
programme. By scanning large numbers of flat-racing Thoroughbreds in two
training yards in Newmarket, we know that the amount of heart muscle,
and the width and area of the left heart chamber increase dramatically
with training. Increased heart volume allows the heart to hold more
blood at rest and during exercise, whilst increased muscle also allows
the more powerful "fit" heart to eject more of this blood with
each beat.
How to measure
heart rate during exercise:
Heart rate monitors:
These monitors detect the electrical signals that instruct the heart to
beat. As each electrical signal is accompanied by a physical contraction
of the heart, they are accurate monitors of heart rate. The horse wears
flat electrodes and a transmitter device under the saddle. The heart
rate is transmitted to a watch monitor worn by the rider. These monitors
can be of varying degrees of complexity and expense: All of them give an
instantaneous heart rate, the most expensive units allow accumulated
heart rate information from one or more training sessions to be
downloaded into a personal computer.
What can my horse's
heart rate tell me?
Increased stroke volume caused by training means that the athletes heart
beats less frequently at all times, including during periods of rest.
Because of the greater volume ejected from the "fit" heart
with each beat, cardiac output remains the same as in the untrained
state, although the heart beats less often. So it should be possible to
see a difference in your horse's heart rate at rest before and after
training. Indeed the true resting heart rates of conditioned racehorses
are often at, or even below, 30 beats per minute. In human athletes the
effect of training on resting heart rate is marked. For example, the
average heart rate of the competitors in the Tour de France cycle race
was below 40 beats per minute compared to the average for a normal man
of 75 per minute. In horses, with a much lower resting heart rate to
start, not only is the effect much less marked, but their character
often masks our ability to detect it. In horses heart rate is greatly
influenced by stress and anxiety. The very act of measuring heart rate
is often sufficient to increase it by 3 - 4 beats per minute, enough to
mask any changes induced by training.
During exercise
Heart rate during exercise is an invaluable monitor of the
cardiovascular system, fitness and performance. The speed at which heart
rate recovers after work also reflects cardiovascular fitness. During
any exercise session the heart rate can be used to determine when the
horse is sufficiently recovered to gallop or canter again. Heart rate
monitoring also allows us to train specifically for endurance or speed.
This is the basis of interval training, a technique that has been
adopted almost universally in human athletics. Since the advent of
interval training, records in every human track event have been smashed,
as athletes benefited from optimal cardiovascular conditioning. Whilst
breaking records may not be goal of most of this article's readers, it
is important to appreciate that fatigue is the most common cause of
injury in equine athletes. A tired horse will lose his action and be
prone to tendon, joint and ligament strain. When any horse's heart rate
exceeds 200 beats per minute and approaches maximal heart rate, the
horse will fatigue within a short time. No matter how much he is urged
the horse cannot give any more.The table shows the average heart rate
and respiratory rates of conditioned Thoroughbreds undergoing treadmill
exercise at gradually increasing speed. You can see how heart rate
increases as the workload goes up to the next speed.
| Gait/Speed |
Average
Heart Rate (b.p.m.) |
| Rest |
35 |
| Walk |
75 |
| Trot |
127 |
| Canter @
7m/s (16 mph) |
170 |
| Canter @
8m/s (18 mph) |
182 |
| Canter @
10m/s (22 mph) |
201 |
| Canter @
11m/s (25 mph) |
219 |
| Canter @
12m/s (27 mph) |
224 |
| Walk - 1
min recovery |
106 |
| Walk - 2
min recovery |
89 |
When an individual's
heart rate is monitored over time and over a standard piece of work, the
rider knows exactly how hard the horse is working on any given day. When
the workload is standardised, individual horses can be compared with
each other to assess their fitness and ability. In every sport, serious
athletes and trainers use heart rate during exercise to devise training
plans and monitor progress. Unfortunately in the equine world we are
still along way behind. Instead of using the heart rate of our horses,
to gauge their effort and their recovery, we guess by relying on how the
horse feels, whether he pulls, or needs encouragement, or how long he
blows once he stops. Unfortunately these are not very accurate methods.
In the past the equipment available to monitor heart rate during
exercise was unsuitable for horses, so there was no alternative than to
follow the traditional methods. Now we no longer have this excuse,
affordable equipment is becoming readily available.
Heart rate
monitors:
What are they?
These monitors detect the electrical signals that instruct the heart
to beat, each electrical signal triggers a physical contraction of the
heart. To pick up this pulse of electricity two conducting electrodes
must be placed on the horse's chest above and below the heart. When the
horse exercises, the sweat produced is sufficient to maintain electrical
contact between the electrodes and the skin. The horse wears the
electrodes and a small transmitter device attached to the saddle and
girth. The only requirement is to keep the sensors pressed against the
skin and keep movement to a minimum. Each time the heart beats, the
transmitter sends a radio pulse to a watch receiver worn by the rider.
The receiver counts the pulses and calculates heart rate and displays
it. As most readers will be aware, heart rate monitors are widely used
by human athletes and the same watch computers are perfectly suitable
for horses. The only adaptation needed to make the units work, is to
alter the electrodes. In the human breast strap, the two electrodes are
positioned 4 inches apart which is sufficient to span the small human
heart. However the equine heart is much bigger, so our electrodes must
be much further apart.
What do they cost?
A basic horse heart monitor starts at around $150.00. The monitors
can be of varying complexity and expense: All of them give an
instantaneous heart rate, and others allow the rider to pre-program
heart rate zones and provide audible alarms and timers to assist with
monitoring training. The most sophisticated and expensive units allow
accumulated heart rate information from one or more training session to
be downloaded into a personal computer. It is worth remembering that the
transmitters work for all the monitors, as a result the rider can
upgrade from a basic to a more sophisticated system, or buy adapted
electrodes and use the watch he uses on himself on his horse!
How easy are heart
rate monitors to use?
Heart rate monitors with a reliable system of electrodes are
extremely easy to use. The electrodes under the saddle are well
tolerated by the horse and cause no irritation. Batteries can be readily
replaced in both the watches and the transmitters. Battery life is up to
5,000 hours
Why is heart rate
important?
Resting heart rate is between 28 and 36 beats per minute in Thoroughbred
horses. Maximum heart rate during peak exercise in the Thoroughbred lies
between 225 and 240 beats per minute. The huge range of heart rates
available to the equine athlete is the most important mechanism for
increasing the output of the heart during exercise.
Heart rate
monitors:
How do they help endurance horses?
At low heart rates ( <140 beats per minute ) the heart is able to
supply the muscles with all the oxygen that they need. The muscles are
able to burn fuel very efficiently. This is called aerobic exercise.
This type of exercise can be carried out for very long periods without
fatigue. When a horse stops after aerobic work his heart rate rapidly
returns to normal and he will hardly blow. In endurance competitions the
goal is to keep the horse exercising at this level throughout. This
allows the very long distances to be covered most efficiently. Keeping
heart rate within this aerobic zone ensures rapid heart rate recovery
when exercise stops. The end result being that the horse exercises
happily for longer and is rapidly released from veterinary checks. For
this reason the most successful endurance horse is rarely the fastest. A
canny competitor will use a heart rate monitor to keep his horse in the
aerobic zone and make sure he's quickest out of the vet gate!
At higher heart rates
(up to 240 beats per minute) the heart can no longer supply the muscles
with all the oxygen that they need. Fuel burning in the muscles is less
efficient and a substance called lactate builds up in the blood. The
heart rate at which lactate begins to accumulate is known as lactate
threshold. Once this heart rate is attained, any increase in speed, or
effort, results in more lactate entering the blood and is called
anaerobic exercise. High heart rates and the build up of lactate rapidly
cause fatigue, so anaerobic exercise cannot be continued for long
periods. Galloping, sprinting and hill work all increase heart rate and
cause lactate production. Heart rate recovery following anaerobic
exercise is much slower, the body is in oxygen debt and the accumulated
lactate must be disposed of; this process takes time. This means a long
wait before an endurance horse is released from a veterinary gate:
Plenty of time for the rider to watch, whilst slower horses arriving
much later, wave a cheery "Goodbye" as they leave. During
anaerobic exercise the muscles use more fuel and generate more heat.
This is not a problem during a 3-minute flat race, but in a 100-mile
endurance ride, a horse that spends long periods above his lactate
threshold will deplete his energy reserves and dehydrate long before his
event is completed.
Because of differences
in the ability of muscles to use oxygen and in the heart's ability to
supply oxygen, each horse's heart rate response to exercise is
different. Each horse also has a different lactate threshold. A rider
who uses a heart rate monitor during competition can ensure his horse
stays comfortably within his aerobic zone whatever the terrain or
climate. At home, the monitor is equally useful for interval training:
Multiple short periods (1 - 5 minutes) of work at speeds, or gradients,
inducing higher heart rates (anaerobic exercise) separated by periods of
work at lower intensity, are very useful for improving the body's
ability to handle lactate. This means as the horse gets fitter, he can
go much faster with a lower heart rate and minimal accumulation of
lactate. In other words, the rider has created the ultimate endurance
horse.
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